Supported by the National Natural Science Foundation of China (No.30470344 and 30670088), Key Technology Project of Hubei Province (No.2006AA305A04) and Commonweal Project of State Oceanic Administration of China (No.200705014),ACKNOWLEDGMENT We thank SHI Zhengli and CHEN Xinwen for creative advice on experimental design and ZHANG Jianhong for providing detailed technological support. We would also like to thank HOU Kun, SUN Qian and YU Rong for the sampling of the lakes.