The association between plants with long corolla tubes and pollinators with long tongues has evolved repeatedly in diverse pollination systems. However, the morphogenesis of long corolla tubes involved in diverse plant groups remains largely unexplored, limiting our understanding of the evolution and maintenance of long corolla tubes. Corolla tubes of Himalayan Pedicularis species exhibit striking length diversity, ranging from less than 10 mm to more than 120 mm. We investigated the role of cell division and cell expansion in determining the corolla tube morphology in eight Pedicularis specie...