This work was supported by the National Natural Science Foundation of China under Grants 61972290 and 61972173 , the National Key R&D Program of China under Grant 2018YFC1604000 , the Fundamental Research Funds for the Central Universities (No. CCNU22QN015 ), and Wuhan knowledge innovation project (No. 2022010801010278 ). This work was also supported by the Natural Sciences and Engineering Research Council (NSERC) of Canada , an NSERC CREATE award in ADERSIM, 4 4 the York Research Chairs (YRC) program and an ORF-RE (Ontario Research Fund-Research Excellence) award in BRAIN Alliance. 5 5 The Authors thank the anonymous reviewers and the associate editor for their insightful comments.