We thank middle-school student J.-Y. Huang, lab members Z.-B. Chen, L.-B. Jia, B. Wang, S.-S. Wang, and B. Wu for field assistance and Z.-H. Huang, S.-G. Sun, Z.-Y. Tong, and Y.-Z. Xiong for suggestions for methodologies and statistical analysis; Z.-D. Fang and staff of Shangri-La Botanical Garden for logistical support; S. Corbet, S. Armbruster, S. Renner, J. Thomson, and three anonymous reviewers for sharing ideas and valuable suggestions for this work; and S. Corbet for correcting the English of the manuscript. This work was supported by the National Natural Science Foundation of China (grant no. 31730012 ) and Guizhou Science and Technology Department Basic Research Foundation ( 2019/1237 ).