1This work was supported by the National Natural Science Foundation of China (grant nos. 31670332 and 31600190), Fundamental Research Funds for the Central Universities (CCNU18ZDPY05 and CCNU19QD013), and the Hubei Provincial Natural Science Foundation (grant no. 2013CFB211). 2These authors contributed equally to the article. 3Author for contact: bsqiu@mail.ccnu.edu.cn. 4Senior author. The author responsible for distribution of materials integral to the findings presented in this article in accordance with the policy described in the Instructions for Authors (www.plantphysiol.org) is: Bao-Sheng Qiu (bsqiu@mail.ccnu.edu.cn). B.-S.Q. and G.-Z.D. conceived and supervised the study; H.-F.X., G.-Z.D., D.-M.Y., and W.-Y.S. conducted the major analyses in this work; J.-L.S. was engaged in the bioinformatics analysis; H.-F.X., G.-Z.D., H.S., and B.-S.Q. analyzed the data and wrote the article. www.plantphysiol.org/cgi/doi/10.1104/pp.19.01149
