We thank F. Wang, Y. Li and X. Shang for assistances with seed collection. M. Steele recognizes the support from the H. Fenner Endowment of Wilkes University , The Howard Hughes Medical Institute , and a Bullard Fellowship from Harvard Forest, Harvard University . H. Zhang thanks the China Scholarship Council and Wilkes University for supporting a visiting scholar position at Wilkes University ([2011]3008). This work was partially supported by the National Natural Science Foundation of China ( 31372212 , 31172102 , 30930016 ) and the State Basic Research Programme of China ( 2007BC109102 ).