Conceptualization, R.P. and K.L.; methodology, R.P., K.G., Y.Z., W.Z., X.G. and L.H.; formal analysis, R.P., K.G., Y.Z., X.Z. and K.L.; investigation, K.G., Y.Z., W.Z., X.G. and L.H.; resources, K.G., Y.Z., K.L., H.Y., H.H., J.P. and R.P.; writing—original draft preparation, R.P., K.G., Y.Z. and X.Z.; writing—review and editing, all authors; supervision, R.P., K.L., H.Y., H.H. and J.P.; funding acquisition, R.P., K.L., H.H. and J.P. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Natural Science Foundation of China (Grant No.31401762, No.31672355), the National Key Research and Development Program of China (Grant No.2017YFD0200400), Wuhan Youth Science and Technology Plan Project of China (Grant No. 2017050304010320), and Fund of Hubei Key Laboratory of Genetic Regulation and Integrative Biology (Grant No. GRIB202210).
