Acknowledgements We thank Yihao Zhang, Jia Li, and Minghui Wang for helping with the experimental setup. We would like to give our thanks to anonymous reviewers for their evaluable comments on the earlier version of this manuscript. Funding This study was supported by the State Key Laboratory of Integrated Management of Pest Insects and Rodents (ChineseIPM1703), the Natural Science Foundation of Jiangxi Province (20161ACB20017, 20171BBF60073), and the Natural Science Foundation of China (31760156 and 31772471).