We thank Professor Scott L. O'Neill (Monash University, Australia) for providing Dmel Mel flies, Professor Dahua Chen (Institute of Zoology, Chinese Academy of Sciences) for providing bamGal4 fly line, and Beijing Genomics Institute (BGI, Shenzhen, China) for technical assistance in identification and quantitation of phosphopeptides and some data analyses. This work was supported by the National Natural Science Foundation of China (no. 31672352, 31872288). w