Conceptualization, Y.L. and X.Y.; methodology, H.Y.; software, Y.L., Z.C. and Z.Z.; validation, Y.M. and S.W.; formal analysis, J.C. and C.Y.; investigation, H.Y.; resources, X.Y.; data curation, J.W.; writing—original draft preparation, Y.L.; writing—review and editing, J.C. and C.Y.; visualization, J.C.; supervision, X.Y.; project administration, Y.L. and X.Y.; funding acquisition, X.Y. All authors have read and agreed to the published version of the manuscript. This study was financially supported by grants from the Key Project of Key Research and Development Program of Ningxia Hui Autonomous Region, China (2018BEG02001), the National Natural Science Foundations of China (32160344), the 2022 Central Finance Forestry Science and Technology Promotion Demonstration Project ([2022]ZY09), and Investigation, Collection and Conservation of Wild plant Germplasm Resources of the National Major Science and Technology Infrastructure “Southwest China Wild Germplasm Resources Bank” (WGB-1514, 1605, 2103).