1This work was supported by The National Natural Sciences Foundation of China (grant nos. 31271637 and 31571572) and The Project of National Research and Development of China (grant no. 2016YFD0100202). 2These authors contributed equally to the article. 3Address correspondence to renfeng@mail.ccnu.edu.cn. 4Senior author. The author responsible for distribution of materials integral to the findings presented in this article in accordance with the policy described in the Instructions for Authors (www.plantphysiol.org) is: Feng Ren (renfeng@mail.ccnu.edu.cn). F.R., K.-L.H., and G.-J.M. designed the study; K.-L.H., G.-J.M., M.-L.Z., H.X., H.W., C.-Z.Z., C.-S.L., H.-X.J., and F.R. performed the experiments; K.-L.H., G.-J.M., X.-B.L., and F.R. analyzed the data; K.-L.H., L.C., J.O.K., and F.R. wrote the article. www.plantphysiol.org/cgi/doi/10.1104/pp.17.01713
