1This work was supported by the National Natural Sciences Foundation of China (grant nos. 31070281 and 31500987). 2These authors contributed equally to the article. 3Author for contact: xbli@mail.ccnu.edu.cn. 4Senior author. The author responsible for distribution of materials integral to the findings presented in this article in accordance with the policy described in the Instructions for Authors (www.plantcell.org) is: Xue-Bao Li (xbli@mail.ccnu.edu.cn). X.-B.L., Y.L., and L.L. conceived and designed the research; Y.L., L.L., Y.W., Y.-C.W., N.-N.W., R.L., and Y.-W.W. performed the experiments; Y.L., L.L., and X.-B.L. analyzed the data and wrote the article. www.plantphysiol.org/cgi/doi/10.1104/pp.20.00297
